Strategy for fighting both of Natla's forms is detailed in the walkthrough for the level. These screenshots are meant to illustrate certain points made there.
Note that I had to use screenshots from several different battles in order to cover everything. So you may notice that the ammo counts, health status and positioning aren't necessarily sequential from one shot to the next.
Fire at Natla constantly as you dodge her fireball attacks.
Keep an eye on her health and rage bars at the top right corner of the screen. When the rage bar maxes out, both bars will glow red, as shown above.
She'll let out a shriek, which is visible as stylized sound waves emanating from her head, and charge at Lara.
Maintain a target lock as you run to meet her. But just before you reach her, roll forward. As long as the enemy is enraged and charging, you have a target lock, Lara is moving forward before you begin rolling, and you roll soon enough that you don't smack into Natla, this will work as an adrenaline dodge.
NOTE: You can also perform an effective dodge by rolling backward, away from Natla, as long as you have room enough to maneuver.
You'll see two separate targeting reticules converge on Natla. . . .
. . . as Lara leaps over her foe.
When the reticules overlap, immediately shoot.
If your shot connects, Natla will be momentarily stunned, and she'll slump over, revealing the green spot on her back. Switch to more powerful weapons if possible and shoot the glowing green spot for as long as you can. This is the only way to injure her.
After a few moments, she'll pull herself together and start the fireball attacks again. Repeat the whole process until you reduce her health to zero.
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